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Within each papilla are hundreds of taste buds, the organ of taste transduction.
It is the primary organ of taste (gustation), as much of the upper surface of the tongue is covered in papillae and taste buds.
They manifest themselves in the contact of substances with the organs of taste, touch, and smell, or otherwise affect the living organism, as in the manner of medicines and poisons.
Karelius coped by employing a technique he had first developed at university and perfected in the army: that of chewing and swallowing whilst managing to divorce his mind completely from the organs of taste.
Tasting means a being affected by what can be tasted as such; hence the organ of taste must be liquefied, and so to start with must be non-liquid but capable of liquefaction without loss of its distinctive nature.